The sub-captain on the Bilkis. He is Tiki's older brother. Mario believes in the concept of love and feels the world is more than just DNA. He felt close to his old Register, but feels closer to Beatrice.
Mario takes over as captain when Hiroto starts to halucinate and falls ill. He awakens memories that Beatrice thought she'd 'deleted' from her brain. As a child, he once ate a register's pill. Of course, not being a register, it had no effect.
Mario is good at lying and leading. He was 2nd only to Jean in his class at school, and they are 'rivals' now. When Mir creates a teleportation system, he uses it to trasport to Jean's ship, the Youki.
Just before dying, Mario says he's going to see love for himself, and Jean says, "Show me Mario".