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Thundercracker (サンダークラッカー)Thundercracker claims the skies as his arena. He looks down in contempt on those who cannot leave the ground — he considers them lesser beings. He will often attack the flightless beings below him for no other reason than to satisfy his scorn.
His superiority in this realm is the only thing he is sure of. Thundercracker is not entirely committed to the Decepticon cause, and even has some seemingly contradictory sympathy for the humans they terrorize. Fear of retribution from Megatron and the influence of his peers is usually enough to bury these doubts. Regardless, he needs constant reassurance from Skywarp and Starscream (which occasionally take the form of threats) that he's where he needs to be. Sometimes, his heart's just not in the job.
In the early days of the Decepticon (Destron in Japanese version) rebellion, Thundercracker and Skywarp both faced the powerful protectors of Cybertron's cities, the Guardian robots (Japanese version is known as: GADEP - Groundling Auto-Destruct Powerbroker). According to Thundercracker, he was often not very successful. Thundercracker was one of Megatron's elite warriors, participating in the attack on the Ark (Iacon Bridge in Japanese version) and joining the rest of the Earth-bound Transformers in their several-million-year nap under an inactive volcano. He awoke when the rest of the Decepticons did, and took part in their attempts to gather energon and escape the planet Earth.
(Source: Transformers Wikia, edited)